I still don't understand how burials indicate how it was Homo Nadeli who made the marks. All it would take is a single human (maybe some teenagers) wandering down one afternoon on a lark and making the marks.
Having read his "Almost Human" book... these are not caves you casually enter. The passages to get to the Naledi burial chambers are impassable to adult males and most women, and are very deep and far in the cave system. He specifically recruited petite women with caving experience to work on his projects.
"A portion of the cave, used by the excavation team en route to the Dinaledi Chamber, is called "Superman's Crawl" because most people can fit through only by holding one arm tightly against the body and extending the other above the head, in the manner of Superman in flight"
That makes it even more likely it could have been humans, doesn’t it? Over a long period of time, probably lots of humans wandered deeper into the cave.